PHYSOPTYCHIS Boiss., Fl. Orient. 1: 260 (1867).

Herbs perennial, cespitose, canescent, with many-branched surculose to compact caudex the branches of which covered with dry persistent leaves of previous seasons and terminated in rosettes. Trichomes short stalked to subsessile, 8–12(–16)-rayed, subdendritic stellate with non-appressed soft rays, simple trichomes absent. Multicellular glands absent. Stems erect to ascending, simple, leafy, not spiny. Basal leaves short petiolate, rosulate, densely pubescent, simple, entire; cauline leaves subsessile or sessile, cuneate to attenuate, not auriculate at base, entire. Racemes several flowered, ebracteate, corymbose, slightly not elongated in fruit; rachis straight, densely pubescent; fruiting pedicels divaricate, straight, densely pubescent, persistent. Sepals oblong, free, deciduous, pubescent, erect, unequal, base of lateral pair saccate; petals yellow, erect at base with flaring blade, longer than sepals; blade obovate, apex obtuse; claw strongly differentiated from blade, subequaling sepals, glabrous, unappendaged, entire; stamens 6, slightly exserted, erect or spreading, strongly tetradynamous; filaments wingless, glabrous, free, lateral pair unilaterally toothed above base; anthers oblong, not apiculate; nectar glands 4, lateral, 1 on each side of lateral stamens, median nectaries absent; ovules 4–8 per ovary; placentation parietal. Fruit indehiscent, bladderlike silicles, globose to ovoid, with rounded to acute apex, terete, strongly inflated, unsegmented; valves thin papery, veinless, moderately to sparsely pubescent outside, glabrous inside, not keeled, smooth, wingless, unappendaged; gynophore obsolete or to 0.5 mm long; replum rounded, concealed by valve margins; septum complete, membranous, veinless; style 1–4 mm long, filiform, pubescent on proximal half, caducous; stigma capitate, entire, unappendaged. Seeds biseriate, narrowly winged or margined all around, ovate to suborbicular, flattened, on funicles adnate to septum; seed coat not mucilaginous when wetted; cotyledons accumbent.