HORMATHOPHYLLA Cullen & T.R.Dudley, Feddes Repert. 71 : 225 (1965).

Subshrubs or suffruticose, sometimes perennial herbs with woody caudex, canescent. Trichomes sessile or subsessile, 8–24-rayed stellate with soft or rigid appressed rays, without webbing between the rays, sometimes with fewer rays on pedicels and sepals than on leaves. Multicellular glands absent. Stems erect to ascending, simple or branched above, leafy, not or rarely spiny. Basal and lowermost cauline petiolate, rosulate or not, simple, entire; cauline leaves subsessile and cuneate to attenuate and not auriculate at base, entire. Racemes several to many flowered, ebracteate throughout or only basally, corymbose or lax, sometimes corymbs or panicles, slightly elongated in fruit; rachis straight; fruiting pedicels ascending to divaricate, persistent. Sepals ovate to oblong, free, deciduous, ascending to spreading, equal or unequal, base of lateral pair saccate or not; petals white or rarely pink, erect at base with flaring blade, longer than sepals; blade obovate to spatulate, apex obtuse or truncate; claw slightly or strongly differentiated from blade, shorter than sepals, glabrous, unappendaged, entire; stamens 6, slightly exserted included, erect, slightly tetradynamous; filaments wingless, toothless, unappendaged, free, glabrous; anthers ovate, minutely apiculate or not; nectar glands 4, lateral, 1 on each side of lateral stamens, median glands absent; ovules 4–8 per ovary; placentation parietal. Fruit dehiscent, capsular silicles, obovate, orbicular, elliptic, or cochleariform, strongly latiseptate, not inflated, unsegmented; valves papery, reticulate veined, glabrous or rarely pubescent, not keeled, smooth, wingless, unappendaged; gynophore obsolete; replum rounded, visible; septum complete, perforated, reduced to a rim, or absent, membranous, thickened, veinless or without a midvein or 1–3 longitudinal veins, rarely densely fibered or veined or forming transverse partitions between seeds; style 0.2–2.5 mm long, filiform, persistent; stigma capitate, entire or slightly 2-lobed, the lobes not connivent, free, not decurrent, opposite replum, unappendaged. Seeds biseriate or aseriate, winged all around or distally, oblong, ovate, or orbicular, flattened; seed coat mucilaginous or not when wetted; cotyledons accumbent.