BERTEROA DC., Mém. Mus. Hist. Nat. 7: 232 (1821).

Herbs annual or perennial with caudex. Trichomes sessile or subsessile, rigid or soft, 3–8-rayed stellate mixed with simple, malpighiaceous, and stalked forked ones on various parts. Multicellular glands absent. Stems erect to ascending, simple or branched basally and/or apically. Basal leaves petiolate, not rosulate, simple, entire or dentate to sinuate; cauline leaves sessile or subsessile, cuneate to attenuate, not auriculate at base, often entire. Racemes many flowered, ebracteate, corymbose, elongated considerably in fruit; rachis straight; fruiting pedicels slender, erect to divaricate, straight, pubescent. Sepals oblong, free, deciduous, suberect to spreading, pubescent, equal, base of inner pair not saccate; petals white or rarely yellow, erect at base with flaring blade, much longer than sepals; blade obcordate, apex deeply 2-fid to about middle; claw short, obscurely differentiated from blade, glabrous, unappendaged, entire; stamens 6, slightly exserted, erect, tetradynamous; filaments filiform, glabrous, free, wingless, those of median pairs slightly or strongly flattened at base, unappendaged or laterally 1-toothed, lateral pair with a basal, adaxial toothlike appendage; anthers oblong, sagittate at base, not apiculate at apex; nectar glands 4, 1 on each side of lateral stamen, median nectaries absent; ovules 4–16 per ovary; placentation parietal. Fruit dehiscent capsular silicles, oblong, elliptic, ovate, obovate, or orbicular, slightly to strongly flattened and latiseptate, sometimes slightly inflated, sessile, unsegmented; valves papery, veinless or with an obscure midvein, stellate hairy or glabrous, smooth, wingless, unappendaged; gynophore absent; replum rounded, visible; septum complete, membranous, veinless; style distinct 0.5–4 mm, cylindric, persistent, glabrous or sparsely pubescent; stigma capitate, obscurely 2-lobed, unappendaged. Seeds biseriate, winged, margined, or wingless, suborbicular to ovate or lenticular, plump or flattened; seed coat minutely reticulate, not mucilaginous when wetted; cotyledons accumbent.